Stuart Lancaster starts his reign as England’s full time coach on Saturday in the opening Test against South Africa in Durban.
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Lancaster did well as England’s interim coach during the RBS Six Nations tournament, where England only lost once, and he has since been appointed full time coach and he will test himself against the Springboks in a three Test series.
He has opted to make four changes to the team that faced Ireland in their final RBS Six Nations game.
Marler will start at loosehead prop after impressing for the Harlequins this season, while Johnson gets the nod with Tom Croft out through injury.
There is a change at full-back with Mike Brown coming in with Ben Foden moving on to the wing due to an injury to David Strettle, while there is a new half-back partnership with Owen Farrell lining up at fly-half alongside Ben Youngs.
New Springboks coach Heyneke Meyer will oversee his first Test match this weekend and he has opted to put his own spin on the side.
Lock forwards Juandre Kruger and Eben Etzebeth will both start at Durban’s Kings Parks, while flanker Marcell Coetzee has also been rewarded with his first start.
There will be a fourth debutant in Durban with prop forward Coenie Oosthuizen looking to get on from the replacements bench.
The Springboks can be backed at 4/11 with Bet365
to win the game, while England are a 9/4 shot. The draw is available at 22/1.
England have been handed a seven point start on the handicap market and can be backed at 10/11 with BetVictor
, while the Springboks -7 are the same price with Blue Square